From Koko.org:
During the course of the study, Koko has advanced further with language than any other non-human. Koko has a working vocabulary of over 1000 signs. Koko understands approximately 2,000 words of spoken English. Koko initiates the majority of conversations with her human companions and typically constructs statements averaging three to six words. Koko has a tested IQ of between 70 and 95 on a human scale, where 100 is considered "normal."
In addition to intensive studies of vocabulary acquisition, the project has investigated spontaneous gorilla language use. This involves the study of innovative linguistic strategies, invention of new signs and compound words, simultaneous signing, self-directed signing, displacement, prevarication, reference to time and emotional states, gestural modulation, metaphorical word use, humor, definition, argument, insult, threat, fantasy play, storytelling and moral judgment. The depth and variety of gorilla language use has significantly exceeded initial expectations. Indeed, evidence has been found for the existence, in less developed form, of almost every aspect of human behavior